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It doesn't run Adsense so it's not just clicking, people buy from ugly sites too.
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When I subscribed to Connected Internet's RSS feed it had a lot less adds. I haven't unsubscribed but I doubt I would actually subscribe anymore. Of course I'm not a good customer because I don't remember ever clicking on an add here.
Maybe revenue is maximised by annoying readers and the lost traffic is worth it. Most likely there is a balance and you need to measure long term trends as well to find the optimum annoyance.
As far as I understand lots of sites target ads not on regular readers but on those that come from search. Searchers couldn't care less about becoming readers they want quick answer and to jump further.
That's why some type of sites require hundreds of thousands visitors to become profitable and some bring money with relatively small traffic.
I think every good site can be profitable but not every profitable site can be good.
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When you change that layout, and in particular 'ugly it up' such that it becomes even more prominent perhaps it draws the eyes of those users. I wonder if over time that uptick would level off to the click rate you're currently used to as people get used to those placements.
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Cheers, Jon
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Great article, thanks for the advice will definitely look at it for my next project.
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Great blog post, thanks.